SAN FRANCISCO • The president of Stanford University said Wednesday that he would resign, citing an independent review that cleared him of research misconduct but found flaws in some papers he wrote.

Marc Tessier-Lavigne said in a statement to students and staff that he would step down Aug. 31.

The resignation comes after the board of trustees launched a review in December following allegations he engaged in fraud and other unethical conduct related to his research and papers.

Tessier-Lavigne, a neuroscientist, says he “never submitted a scientific paper without firmly believing that the data were correct and accurately presented.” But he says he should have been more diligent in seeking corrections regarding his work.

The review assessed 12 papers that Tessier-Lavigne worked on, five of them in which he was the principal author. He said he was aware of issues with four of the five papers but acknowledged taking “insufficient” steps to deal with the issues. He said he’ll retract three of the papers and correct two.

The papers were published before Tessier-Lavigne became Stanford president.

Misconduct allegations about the work were aired on PubPeer, a website where members of the scientific community can discuss research papers, the report stated.

The panel cleared him of the most serious allegations, that a 2009 paper published in the scientific journal Nature was the subject of a fraud investigation and that fraud was found.

There was no investigation and no fraud discovered, the panel ruled. The paper proposed a model of neurodegeneration, which could have great potential for Alzheimer’s disease research and therapy, the panel wrote in its report.

But the panel also concluded the paper had multiple problems, including a lack of rigor in its development and that the research that went into the paper and its presentation contained “various errors and shortcomings.” The panel did not find evidence that Tessier-Lavigne was aware of the lack of rigor.